Sollux, Mituna, Rufioh, Possibly the Serkets (we aren’t sure about the vision eightfold), and maybe Eridan if you subscribe to the theory that his gills are deformed

Vriska

Jade’s inverse is a Seer of Time which she acted like at the beginning of the comic always knowing when things would happen.

(Vriska)’s inverse is Page of Void which she acted like when she mellowed out and Vriska’s piss attitude towards pages is definitely not part of the reason she was so mean to her.

That was obviously Karkat’s opinion since more than one of the trolls is a mutatant

He got it here, when Bro cut it

I’d recommend checking out the wiki page

Mage of Time

I’d think only Derse dreamers can go grimdark since they have a closer tie to the horrorterrors, but that’s up to speculation

She gets ko’ed

because who doesn’t love some alien tiddies???